Venation: - both cells divided - FW quadrifid - three anal veins on HW - FW (R5, (R4, R3)) staked, sometimes from cell, sometimes as (R2 (R5, (R4, R3))). - FW A1 forked at base - HW M1 and R stalked Male antennae usually bipectinate only over the basal half to 2/3 Tympanum absent